TX2i custom board pcie "Response decoding error"

Autonomous Machines Jetson & Embedded Systems Jetson TX2
1

R32.3.1

After forcing a rescan of the PCI buses

[  102.011086] pci_bus 0000:01: busn_res: [bus 01] end is updated to 01
[  102.011100] tegra-pcie 10003000.pcie-controller: PCIE: Response decoding error, signature: 10010001
[  102.020355] pci_bus 0000:02: busn_res: [bus 02] end is updated to 02
[  102.020359] tegra-pcie 10003000.pcie-controller: PCIE: Response decoding error, signature: 10020001

The board is using configuration #1 ODMDATA=9000
There is a 1x1 Intel I210 chip connected to the PEX0 1x4 bus

3

This is the second PCI bus, which is normally connected via M.2 port.
It is unused on our custom board and the hardware engineers tied all the pins to ground, per the OEM guide.